Genus Miarus - Europa

ColeopteraRhynchophoraCurculionidaeMecininae
By Roberto Caldara (sligthly modified by Arved Lompe)
Caldara, R. (2001): Phylogenetic analysis and higher classification of the tribe Mecinini (Coleoptera: Curculionidae, Curculioninae). Kol. Rundschau 71:171-203.
Caldara, R. (2007): Taxonomy and phylogeny of the species of the weevil genus Miarus Schönherr, 1826. (Coleoptera: Curculionidae, Curculioninae). - Koleopterologische Rundschau, 77:199-248 >>>"F:\taxa\coleo\scans\Caldara2007 Miarus.tif"
#1
→ 2♂: Pygidium simple, without median gibbosity and fovea; ventrite 5 without fovea and teeth; elytra somewhat long [Abb. 1]; body length 1.9-2.5 mm. ♀: Pygidium simple and bursa copulatrix without sclerites.
miarus_afer_hab.gif
Abb. 1
→ 3-♂: Pygidium with more or less pronounced median gibbosity and fovea; ventrite 5 without or with more or less deep fovea and two more or less robust teeth laterally to fovea [Abb. 2]; elytra fairly short to short [Abb. 3]; body length 1.6-3.5 mm. ♀: Pygidium with fovea and bursa copulatrix with two semilunate sclerites.
miarus_campanulae_hlb.jpg
Abb. 2
miarus_monticola_hab.jpg
Abb. 3
#2
afer Daniel, 1912Basal margin of elytra nearly regularly and weakly concave, scales of elytral interstriae nearly completely arranged in single regular rows; in ♂ uncus of protibia small [Abb. 4]; median lobe of aedeagus elongate and narrow, with short apex, endophallus with elongate flagellum and without elongate median ventral sclerite [Abb. 5]. Morocco, Algeria, Tunisia.
miarus_afer_vschn.gif
Abb. 4
miarus_afer.gif
Abb. 5
praecursor Caldara, 2001-Basalrand der Flügeldecken von der Naht bis zum 5. Streifen leicht konvex dann deutlich schräg nach vorne verlaufend. Schuppen der Flügeldeckenstreifen 1-2 reihig angeordnet. ♂: Voderschienen mit großem Spitzendorn. Medianlobus des Aedoeagus kürzer und breiter mit langer Spitze; Endophallus mit kurzem Flagellum und langem, ventralem Sklerit. Basal margin of elytra weakly convex from suture to fifth interstria, then distinctly directed forwards; scales of elytral interstriae arranged in 1-2 irregular rows; in ♂ uncus of protibia robust [Abb. 6]; median lobe of aedeagus shorter and wide, with long apex, endophallus with short flagellum and with elongate median ventral sclerite [Abb. 7]. Greece, Turkey, Syria, Jordan.
miarus_praecursor_vschn.gif
Abb. 6
miarus_praecursor.gif
Abb. 7
#3
→ 4Ventrite 5 of ♂ at most with weak depression, covered with scarce to dense hair-like vestiture (Figs. 34-36).
→ 6-Ventrite 5 of ♂ with fovea moderately shallow to deep, nearly glabrous (Figs. 37-40).
#4
stoeckleini Franz, 1947Body length 1.8-2.3 mm; protibiae short, in male with very robust uncus as long as 1/5 of tibia [Abb. 8]; depression of ventrite 5 of male with scarce vestiture [Abb. 9]; median lobe of aedeagus in dorsal view distinctly wider at base than at apex, in lateral view as thin at base as at apex, with orifice very small and narrow [Abb. 10]; habitus as in [Abb. 11]. Croatia, Macedonia, Greece, Turkey, Armenia, Syria.
miarus_stoeckleini_vschn.gif
Abb. 8
miarus_stoeckleini_abd.gif
Abb. 9
miarus_stoeckleini.gif
Abb. 10
miarus_stoeckleini_hab.gif
Abb. 11
→ 5-Body length 2.0-2.8 mm; protibiae longer, in male with moderately robust uncus as long as 1/8 of tibia; depression of ventrite 5 of male with vestiture denser than at sides [Abb. 12]; median lobe of aedeagus in dorsal view as wide at base as at apex, in lateral view distinctly robust at apex, with orifice larger [Abb. 13].
miarus_rotundicollis_abd.gif
Abb. 12
miarus_rotundicollis.gif
Abb. 13
#5
rotundicollis Desbrochers, 1893Pronotum 1/3 wider than long, usually widest near middle; basal margin of elytra regularly concave [Abb. 14]; ventrite 5 of male without posterolateral teeth; median lobe of aedeagus without distinct ventral tongue [Abb. 13], endophallus [Abb. 15]. Albania, Macedonia, Greece.
miarus_rotundicollis_hab.gif
Abb. 14
miarus_rotundicollis.gif
Abb. 13
miarus_rotundicollis_pc.gif
Abb. 15
hellenicus Dieckmann, 1978-Pronotum 1/2 wider than long, widest at basal 1/3; basal margin of elytra usually weakly convex to fifth interstria then distinctly directed forwards [Abb. 16]; ventrite 5 of male usually with two small posterolateral teeth; median lobe of aedeagus with distinct ventral tongue [Abb. 17]. Bulgaria, Greece.
miarus_hellenicus_hab.gif
Abb. 16
miarus_hellenicus.gif
Abb. 17
#6
→ 7Ventrite 5 of male with fovea moderately shallow and teeth usually moderately robust [Abb. 18].
miarus_abnormis_abd.gif
Abb. 18
→ 11-Ventrite 5 of male with fovea deep and teeth usually very robust [Abb. 19].
miarus_ursinus_abd.gif
Abb. 19
#7
→ 8Ventrite 5 of male with a very small shining median tooth anterior to fovea [Abb. 20].
miarus_araxis_abd.gif
Abb. 20
→ 9-Ventrite 5 of male without tooth anterior to fovea.
#8
araxis Reitter, 1907Scales of dorsal vestiture brown, those covering elytral interstriae recumbent [Abb. 21]; rostrum in lateral view distinctly curved, in female only slightly longer than in male; ventrite 5 of male with shallower fovea; median lobe of aedeagus with tip short, not visible in dorsal view [Abb. 22]. Armenia, Georgia, Turkey.
miarus_araxis_hab.gif
Abb. 21
miarus_araxis.gif
Abb. 22
dentiventris Reitter, 1907-Scales of dorsal vestiture white, those covering elytral interstriae suberect to erect [Abb. 23]; rostrum in lateral view in male weakly curved, in female nearly straight [Abb. 24] and somewhat longer than in male; fovea of ventrite 5 deeper; median lobe of aedeagus with tip long, clearly visible in dorsal view [Abb. 25]. Armenia, Iran.
miarus_dentiventris_hab.gif
Abb. 23
miarus_dentiventris_k.gif
Abb. 24
miarus_dentiventris.gif
Abb. 25
#9
solarii Smreczynski, 1957Rostrum in lateral view distinctly tapered in apical 1/3 [Abb. 26]; body length 2.1-2.6 mm; scales of dorsal vestiture whitish [Abb. 27]. Ukraine, Turkey.
miarus_solarii_k.gif
Abb. 26
miarus_solarii_hab.gif
Abb. 27
→ 10-Rostrum in lateral view nearly of same width from base to apex; body length 2.4-3.0; scales of dorsal vestiture whitish to brown.
#10
monticola Petri, 1912Scales of dorsal vestiture whitish [Abb. 28]; median lobe of aedeagus without flanges at apex laterally [Abb. 29]. Finland, north-western Russia (Karelia, Arkhangel'sk Prov., Leningrad Prov., Kirovsk Prov., Bryansk, Moscow Prov.; ZISP), Ukraine, Poland, Germany, Czech Republic, Slovakia, Austria, Bosnia and Herzegovina, Serbia, Albania, Montenegro, Bulgaria, Greece
miarus_monticola_hab.gif
Abb. 28
miarus_monticola3.gif
Abb. 29
abnormis Solari, 1947-Scales of dorsal vestiture whitish to brown or of both colours intermixed; median lobe of aedeagus with distinct flanges at apex laterally [Abb. 30]. South-eastern Poland, Austria, north-eastern Italy, Slovenia, Croatia, Serbia, Montenegro, Macedonia. Its presence in southern France needs confirmation.
miarus_abnormis2.gif
Abb. 30
#11
→ 12Rostrum in female very long (Rl/Pl > 1.30) and distinctly longer than in male [Abb. 31] [Abb. 32] [Abb. 33].
miarus_banaticus_k.gif
Abb. 31
miarus_longicollis_k.gif
Abb. 32
miarus_abeillei_k.gif
Abb. 33
→ 14-Rostrum in female only slightly longer than in male (Rl/Pl < 1.05).
#12
IongicollisScales of elytral interstriae arranged in 1-2 irregular rows, subrecumbent; median lobe of aedeagus at apex with small auricles (au) and very small mitre (m) [Abb. 34]; habitus of female as in [Abb. 35]. Athos, Greece. Only known from the type locality.
miarus_longicollis.gif
Abb. 34
miarus_longicollis_hab.gif
Abb. 35
→ 13-Scales of elytral interstriae mainly arranged in 3-4 irregular rows, recumbent; median lobe of aedeagus at apex with distinct auricles and mitre.
#13
abeillei Desbrochers, 1893Rostrum nearly straight in lateral view in both sexes [Abb. 33] and less punctate in male; scales of dorsal vestiture whitish to brown; median lobe of aedeagus with auricles distinctly uncinate and without spines [Abb. 36]; habitus of female as in [Abb. 37]. South-eastern France, Switzerland, north-western Italy.
miarus_abeillei_k.gif
Abb. 33
miarus_abeillei.gif
Abb. 36
miarus_abeillei_hab.gif
Abb. 37
banaticus Reitter, 1907-Rostrum moderately curved in lateral view in both sexes [Abb. 31] and more punctate in male; scales of dorsal vestiture usually brown or white and brown intermixed; median lobe of aedeagus with auricles not uncinate but usually with some minute spines [Abb. 38]; habitus of female as in [Abb. 39]. Bosnia and Herzegovina, Romania.
miarus_banaticus_k.gif
Abb. 31
miarus_banaticus.gif
Abb. 38
miarus_banaticus_hab.gif
Abb. 39
#14
→ 15Median lobe of aedeagus with medium sized [Abb. 40] to large auricles [Abb. 41].
miarus_ursinus.gif
Abb. 40
miarus_campanulae3.gif
Abb. 41
→ 17-Median lobe of aedeagus at most with small auricles [Abb. 42] [Abb. 43].
miarus_simplex.gif
Abb. 42
miarus_ajugae3.gif
Abb. 43
#15
campanulae (L., 1767)Ventrite 5 of male with fovea usually very deep and teeth distinctly robust [Abb. 2]; median lobe of aedeagus with auricles large, distinctly uncinate [Abb. 41]; body medium to large sized (length 2.3-3.4 mm); scales of elytral interstriae arranged in more than one irregular rows [Abb. 44]; uncus of protibia in male moderately robust [Abb. 45]. Great Britain, Ireland, Spain, France, Belgium, Holland, Poland, Germany, Hungary, Czech Republic, Slovakia, Austria, peninsular Italy, Denmark, Norway, Sweden, Finland, western Russia.
miarus_campanulae_hlb.jpg
Abb. 2
miarus_campanulae3.gif
Abb. 41
miarus_campanulae_hab.jpg
Abb. 44
miarus_campanulae_vschn.gif
Abb. 45
→ 16-Ventrite 5 of male with fovea usually moderately deep and teeth moderately robust [Abb. 19]; median lobe of aedeagus with auricles smaller, not uncinate to moderately uncinate; body small to medium sized (length 1.6-2.8 mm); scales of elytral interstriae often arranged in single regular rows; uncus of protibia in male small.
miarus_ursinus_abd.gif
Abb. 19
#16
ursinus Abeille, 1906Pronotum moderately convex on disc; median lobe of aedeagus with auricles not uncinate and with tongue distinctly shorter [Abb. 40]. South-western Europe: Portugal, Spain, France (incl. Corsica), Switzerland, peninsular Italy, Croatia..
miarus_ursinus.gif
Abb. 40
maroccanus Solari, 1947-Pronotum usually distinctly convex on disc; median lobe of aedeagus with auricles uncinate and with tongue distinctly longer [Abb. 46]. North-western Africa: Morocco, Algeria..
miarus_maroccanus.gif
Abb. 46
#17
atricolor Morimoto, 1983Median lobe of aedeagus with distinct mitre recurved posteriorly [Abb. 47]. Russia (Far East), Kazakhstan, Kyrgyzstan, China, Japan, Korea.
miarus_atricolor.gif
Abb. 47
→ 18-Median lobe of aedeagus without or with very small mitre.
#18
ajugae (Hbst., 1795)Median lobe of aedeagus with mitre and auricles small but distinct, with anteapical stricture very strong, with tongue larger in basal half and clearly visible in dorsal view [Abb. 43]. France, Switzerland, Poland, Germany, Czech Republic, Slovakia, Hungary, Austria, Italy, Slovenia, Croatia, Bosnia and Herzegovina, Serbia, Montenegro, Macedonia, Armenia, Denmark, Sweden, Norway, Finland, Russia (from North West to Far East), Mongolia, Korea, Syria, Iran, north-western Africa.
miarus_ajugae3.gif
Abb. 43
simplex Solari, 1947-Median lobe of aedeagus without mitre and auricles, with anteapical stricture moderately strong, with tongue smaller in basal half and indistinct in dorsal view [Abb. 42]. Spain, France, northern and central Italy, Serbia, Romania, Bulgaria, European Turkey, Georgia.
miarus_simplex.gif
Abb. 42
Enthaltene Taxaabeillei abnormis afer ajugae araxis atricolor banaticus campanulae dentiventris hellenicus Iongicollis maroccanus monticola praecursor rotundicollis simplex solarii stoeckleini ursinus
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